Ecology                  ABIO 320               Fall 2011


Objectives
Prerequisites
Procedures
Texts
Problem Sets
Project
Grades
Syllabus
Test Grades

      Class Number: 1463

      Time: 11:30 AM - 12:25 PM, MWF

      Location: ES 245

      Instructor: Dr. Thomas Caraco

      Office: Biology 253      Office Hours: Thursday, 3:30 - 5:30 PM

      Email: caraco@albany.edu


Course Objectives:

Ecological theory provides a rigorous conceptual framework for understanding the complexity observed across both natural and managed landscapes, and for constructing scientific analyses of applied problems. The course therefore focuses on the core of this framework, a series of well-defined mathematical models for the dynamics of single populations, and for the growth of ecologically interacting species. This course offers motivated students an appreciation of Theoretical Biology. Furthermore, the curriculum prepares any interested students for graduate study of Ecology.

Students should acquire familiarity with basic ecological concepts, by learning to formulate and explain introductory mathematical models of Ecology. Students should gain an understanding of the mathematical analysis of [1] single-population growth in constant and fluctuating environments, [2] interspecific competition, [3] predator-prey interaction, and [4] advance of infectious disease. Students should understand how predictions deduced from theory guide empirical work in "hypothetico-deductive" science.

Students will demonstrate achievement of these objectives by answering in-class quizzes, by writing two in-class examinations, and by completing a project analyzing one or more questions about population dynamics..

... despite the field's reputation as a soft science, nearly all of biology is now ripe for quantitative analysis ...
   Phillips, R., Quake, S., Physics Today, May 2006.

Students should realize that this course addresses quantitative, organizing concepts fundamental to population biology. Hence the course does not include focused study of envirommental problems or conservation technologies. For an intelligent discussion of of contemporary environmental challenges, see Vermont Law School's Environmental Watch List.


Course Prerequisites:

BIO 212Y, Genetics.

MAT 106, Calculus (or higher) OR PHY 140, Mechanics (or higher).

BIO 212Y introduces students to the language of population genetics, evolution and adaptation - necessary to advance in evolutionary ecology. A calculus course, or physics with calculus, is just as essential for ecology. Students should have some familiarity with derivatives, difference equations, and differential equations. Testing will emphasize quantitative problems.

Students may find several of the links listed on The Calculus Page useful.

Advice/Classroom Procedures:

Class attendance is not mandatory. However, in-class quizzes need not be announced prior to the date administered. Read and follow any University at Albany guidelines for missing class when you feel ill.

If attending class, please arrive on time for lecture, and be quiet when lecture begins. The course objectives are acquired more easily if reading assignments (see below) are completed prior to lecture. A fraction of each student's final grade is earned through following course procedures; points are lost by failing to complete assignments on time and by disruptive behavior in class (talking, sleeping, ...).

Texts:

1. Gotelli, N.J. 2008. A Primer of Ecology, 4th Edition.

Dr. Gotelli's book provides a clear, concise introduction to population dynamics and related ecological models.

2. Alstad, D. 2001. Basic Populus Models of Ecology.

Dr. Alstad's book discusses population dynamics, examines a series of epidemic models, and guides the student's numerical investigation of ecological models. The book serves as a "laboratory manual" for Populus, a useful, free software tool. When downloading Populus you first may need to download and install a virtual running environment; the website offers adequate guidance. An alternative, with which this class has far less experience, is a commercial web site.

The two books overlap strongly in topics addressed, since both cover central issues of Ecology. But the books differ in important ways. For example, Dr. Gotelli's book includes discussion of field studies bearing on ecological theory; Dr. Alstad's book devotes a chapter to epidemics, a major focus of population dynamics. Students might find Dr. Gotelli's book sufficient, since Populus includes "help files," summarizing material in Dr. Alstad's book. Remember that books can be purchased (new or used) and that a limited number of copies may be avialble for semester-long rental.

Problem Sets:

Both texts listed above present problems for solution. To help students meet course objectives, links to a series of problem sets are provided below. Most of the problems require analytical or numerical solution. Solving problems proves the best preparation for quizzes and tests. Think about and solve the problems associated with particular topics as we address the associated text material.

Problem Set 1:   Population estimation, Exponential growth

Problem Set 2:   Geometric-mean growth, Logistic growth

       Answer Sheet:   Problem Set 2

Problem Set 3:   Discrete-time logistic growth, Life tables

       Answer Sheet:   Problem Set 3

Problem Set 4:   Population projection, Reproductive value

Problem Set 5:   Metapopulation dynamics, Interspecific competition

       Answer Sheet:   Problem Set 5

Problem Set 6:   Predator-prey dynamics, SIR epidemic

       Answer Sheet:   Problem Set 6

Whoever despises the high wisdom of mathematics nourishes himself on delusion.
   da Vinci, 1489.

Project:

Each student will complete a project summarizing their analysis of an ecological model. The project may involve (i) development of a new model, (ii) numerical investigation of the dynamics of an existing model (Populus would prove useful here), or a similar exercise. For most students, numerical verification of an ecological model's properties will meet the minimal requirement. The project must be summarized in a ten-page document. The project report is due the last class meeting.

View a one-page discussion of project details.



Grade Determination:

Two (2) tests will be administered. Each test equates with 25% of the course grade (hence, 50% in total).

In-class quizzes, each graded pass/fail, collectively equate to 15% of the course grade.

The project report contributes the 30% of the course grade. Following course procedures (see above) earns the final 5% of the course grade.

Students accumulating 90 or more of the 100 available "points" will earn a final grade of A; most students find this goal challenging.

Each test will be administered approximately seven (7) days after competion of the lectures presenting topics covered by the test.

Test 1 will be presented on Monday, 17 October 2011.

Test 2 will be presented on Friday, 18 November 2011.

The course does not include a final exam.

Test 1: percentile scores.

Test 1 + Test 2: percentile scores; letter grades only approximate.



Syllabus

BIO 320 Syllabus
Topic Reading: Text Pages
Density-Independent Dynamics Gotelli: 2-23
Alstad: 1-15
Density-Dependence: Self Regulation Gotelli: 26-48
Alstad: 16-40
Dynamic Overcompensation and Chaos here
Age-Structured Population Dynamics Gotelli: 50-79
Alstad: 41-58, 66-68
Test 1
Metapopulation Dynamics Gotelli: 82-96
Inter-Specific Competition Gotelli: 100-124
Alstad: 71-90
Predator-Prey Dynamics Gotelli: 126-153
Alstad: 91-103
Infection Dynamics Alstad: 115-133
Test 2
Island Biogeography Gotelli: 156-175
Ecological Succession Gotelli: 180-202
Community Diversity Gotelli: 204-224
Geography of Species Number
The Biosphere: Anthropogenic Impact





Recent Test Scores

Last 4 of ID   Quiz 1   Quiz 2    Test 1   Test 2   Quiz 3   Project



Links:   Instructor    Society for Mathematical Biology    Ecological Society of America


Introduction    Note on Evolution    Chapt 1, Lect 1    Chapt 1, Lect 2   

Geometric Mean Growth     Environmental and Demographic Stochasticity     Chapter 1, Final Points   


Introduction to Biotic Regulation     On Logistic Growth     Continuous-Time Logistic Growth    

Temporal Variation In K     Discrete-Time Self-Regulation     Chaos


On Demography    Net Reproductive Rate     Age-Structured Population Projection   

Local Schedule    Topics List, Chapters 1, 2 & 3    More Review Problems    Empirical Insights


Metapopulation Dynamics


Interspecific Interaction: Introduction     Niches, Invasion and Two-Species Competition

Lotka-Volterra Competition        Competition and Character Divergence


Notes on Predator-Prey 1     Notes on Predator-Prey 2

From Lotka-Volterra to Rosenzweig-MacArthur    

      Indirect Interactions   Trophic Cascades


SI and SIR Epidemics    Pathogen Life Cycles

Chapter 6 Summary   

Practice Problems for Test 2


     Island Biogeography


Ecological Succession     Optional: Markov Chain Model for Succession     An Example    

Species Diversity: Intro     Geography and Diversity


My Project     Population Dynamics Project    
The 95-th Annual Meeting of the Ecological Society of America, August 2010, took "Global Warming: The legacy of our past, the challenge for our future" as its theme. The Recent Advances Lecture was presented by Dr. John P. Holdren, Assistant to the President for Science and Technology, and Director, Office of Science and Technology, Office of the President of the United States. Dr. Holdren's spoke on "Energy, the Environment, and the Millenium Assessment of Human Behavior." You may view a copy of his presentation (pdf)
Ecosystems and Human Impact
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Page last updated December 6, 2011 by Dr. Thomas Caraco.